Ovule is also called integumented magasporangiu, It developes into seed after fertlization in spermatophytes. It occurs singly or in a cluster inside ovary with parenchymatous cushions called placenta
The cones bearing megasporophylls with ovules or megasporangia are called macrosporangiate or female strobili.
The male or female cones or strobili may be borne on the same tree (Pinus). However, in cycas male cones and megasporophylls are borne on different trees. The megaspore mother cell is differentiated from one of the cells of the nucellus. The nucellus is protected by envelopes and the composite structure is called an ovule.